Atlas of Genetics and Cytogenetics in Oncology and Haematology


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List of all cancer genes by chromosome

X Y 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 NA

- chromosome 9 -
SymbolGoldenPath (Mb)LocationDescription
FOXD4 106.235 9p11-q11 forkhead box D4
KANK1 696.896 9p24.3 ankyrin repeat domain 15
SMARCA2 2005.342 9p22.3 SWI/SNF related, matrix associated, actin dependent regulator of chromatin, subfamily a, member 2
SLC1A1 4480.444 9p24 solute carrier family 1 (neuronal/epithelial high affinity glutamate transporter, system Xag), member 1
AK3 4701.158 9p24.1-p24.3 adenylate kinase 3
JAK2 4975.245 9p24 Janus kinase 2 (a protein tyrosine kinase)
JBS 4975.245 11q24.1 Jacobsen syndrome
INSL4 5221.419 9p24 insulin-like 4 (placenta)
RLN2 5289.868 9p24.1 relaxin 2
RMRP 5289.868 9p21-p12 RNA component of mitochondrial RNA processing endoribonuclease
RLN1 5324.969 9p24.1 relaxin 1
CD274 5440.559 9p24 CD274 molecule
PDCD1LG2 5500.570 9p24.2 programmed cell death 1 ligand 2
MLANA 5880.909 9p24.1 melan-A
TPD52L3 6318.375 9p24.1 tumor protein D52-like 3
UHRF2 6403.151 9p24.1 ubiquitin-like, containing PHD and RING finger domains, 2
JMJD2C 6747.654 9p24.1 jumonji domain containing 2C
PTPRD 8304.247 9p23-p24.3 protein tyrosine phosphatase, receptor type, D
TYRP1 12683.449 9p23 tyrosinase-related protein 1
NFIB 14071.848 9p24.1 nuclear factor I/B
PSIP1 15454.066 9p22.3 PC4 and SFRS1 interacting protein 1
SH3GL2 17569.194 9p22 SH3-domain GRB2-like 2
ADAMTSL1 18464.104 9p22.2-p22.1 ADAMTS-like 1
ADFP 19105.760 9p22.1 adipose differentiation-related protein
RPS6 19366.254 9p21 ribosomal protein S6
MLLT3 20334.968 9p22 myeloid/lymphoid or mixed-lineage leukemia (trithorax homolog, Drosophila); translocated to, 3
IFNB1 21067.105 9p21 interferon, beta 1, fibroblast
IFNA17 21217.242 9p22 interferon, alpha 17
IFNA13 21357.424 9p22 interferon, alpha 13
IFNA1 21430.440 9p22 interferon, alpha 1
CRAC1 21471.067 15q13-q21 colorectal adenoma and carcinoma 1
RAB7B 21471.067 1q32 RAB7B, member RAS oncogene family
MTAP 21792.635 9p21 methylthioadenosine phosphorylase
COX1 21792.635 - cytochrome c oxidase I
CDKN2A 21957.752 9p21 cyclin-dependent kinase inhibitor 2A (melanoma, p16, inhibits CDK4)
CDKN2B 21992.906 9p21 cyclin-dependent kinase inhibitor 2B (p15, inhibits CDK4)
TUSC1 25666.394 9p21.2 tumor suppressor candidate 1
PLAA 26894.518 9p21 phospholipase A2-activating protein
TEK 27099.286 9p21 TEK tyrosine kinase, endothelial (venous malformations, multiple cutaneous and mucosal)
C9orf14 27235.683 9p21 chromosome 9 open reading frame 14
ACO1 32374.658 9p22-q32|9p21.1 aconitase 1, soluble
DDX58 32445.301 9p12 DEAD (Asp-Glu-Ala-Asp) box polypeptide 58
TOPORS 32530.543 9p21 topoisomerase I binding, arginine/serine-rich
TAF1L 32619.452 9p21.1 TAF1 RNA polymerase II, TATA box binding protein (TBP)-associated factor, 210kDa-like
APTX 32962.609 9p13.3 aprataxin
DNAJA1 33015.209 9p13-p12 DnaJ (Hsp40) homolog, subfamily A, member 1
B4GALT1 33100.642 9p13 UDP-Gal:betaGlcNAc beta 1,4- galactosyltransferase, polypeptide 1
BAG1 33242.470 9p12 BCL2-associated athanogene
AQP3 33431.162 9p13 aquaporin 3 (Gill blood group)
PTENP1 33665.634 9p21 phosphatase and tensin homolog (mutated in multiple advanced cancers 1), pseudogene 1
PRSS3 33785.559 9p11.2 protease, serine, 3 (mesotrypsin)
UBE2R2 33807.182 9p13.3 ubiquitin-conjugating enzyme E2R 2
UBAP1 34169.011 9p13.3 ubiquitin associated protein 1
NUDT2 34319.504 9p13 nudix (nucleoside diphosphate linked moiety X)-type motif 2
CNTFR 34541.431 9p13 ciliary neurotrophic factor receptor
OPRS1 34624.720 9p13.3 opioid receptor, sigma 1
GALT 34636.635 9p13 galactose-1-phosphate uridylyltransferase
IL11RA 34643.932 9p13 interleukin 11 receptor, alpha
CCL27 34651.893 9p13 chemokine (C-C motif) ligand 27
CCL19 34679.569 9p13 chemokine (C-C motif) ligand 19
CCL21 34699.004 9p13 chemokine (C-C motif) ligand 21
DNAJB5 34979.785 9p13.3 DnaJ (Hsp40) homolog, subfamily B, member 5
VCP 35046.561 9p13.3 valosin-containing protein
FANCG 35063.835 9p13 Fanconi anemia, complementation group G
PIGO 35078.690 9p13.3 phosphatidylinositol glycan anchor biosynthesis, class O
STOML2 35089.893 9p13.1 stomatin (EPB72)-like 2
TESK1 35595.281 9p13 testis-specific kinase 1
CD72 35599.976 9p13.3 CD72 molecule
C9orf100 35649.341 9p13.3 chromosome 9 open reading frame 100
CA9 35663.915 9p13-p12 carbonic anhydrase IX
TLN1 35687.334 9p13 talin 1
NPR2 35782.406 9p21-p12 natriuretic peptide receptor B/guanylate cyclase B (atrionatriuretic peptide receptor B)
C9orf127 35819.513 9p13.3 chromosome 9 open reading frame 127
RECK 36026.916 9p13-p12 reversion-inducing-cysteine-rich protein with kazal motifs
GNE 36204.441 9p13.3 glucosamine (UDP-N-acetyl)-2-epimerase/N-acetylmannosamine kinase
RNF38 36326.401 9p13 ring finger protein 38
MELK 36562.905 9p13.2 maternal embryonic leucine zipper kinase
PAX5 36828.531 9p13 paired box 5
GRHPR 37412.707 9q12 glyoxylate reductase/hydroxypyruvate reductase
RAB1C 37626.933 9p13.2 RAB1C, member RAS oncogene family pseudogene
ALDH1B1 38382.702 9p11.1 aldehyde dehydrogenase 1 family, member B1
IGFBPL1 38398.992 9p13.1 insulin-like growth factor binding protein-like 1
FOXD4L4 69666.445 9q12 forkhead box D4-like 4
FOXD4L3 70107.603 9q21.11 forkhead box D4-like 3
PIP5K1B 70510.436 9q13 phosphatidylinositol-4-phosphate 5-kinase, type I, beta
PRKACG 70817.271 9q13 protein kinase, cAMP-dependent, catalytic, gamma
FXN 70840.164 9q13-q21.1 frataxin
FRA11B 70840.164 11q23.3 fragile site, folic acid type, rare, fra(11)(q23.3)
FRA11F 70840.164 11q14.2 fragile site, aphidicolin type, common, fra(11)(q14.2)
FRA6E 70840.164 6q26 fragile site, aphidicolin type, common, fra(6)(q26)
FRA6F 70840.164 6q21 fragile site, aphidicolin type, common, fra(6)(q21)
FRA9E 70840.164 9q32 fragile site, aphidicolin type, common, fra(9)(q32)
TJP2 70978.909 9q13-q21 tight junction protein 2 (zona occludens 2)
SMC5 72063.698 9q21.11 structural maintenance of chromosomes 5
GDA 73954.113 9q21.13 guanine deaminase
ALDH1A1 74705.408 9q21.13 aldehyde dehydrogenase 1 family, member A1
ANXA1 74956.601 9q12-q21.2|9q12-q21.2 annexin A1
RORB 76302.072 9q22 RAR-related orphan receptor B
TRPM6 76527.231 9q21.13 transient receptor potential cation channel, subfamily M, member 6
GCNT1 78246.402 9q13 glucosaminyl (N-acetyl) transferase 1, core 2 (beta-1,6-N-acetylglucosaminyltransferase)
PCA3 78569.171 9q21-q22 prostate cancer antigen 3
PRUNE2 78628.003 9q21.13 prune homolog 2 (Drosophila)
FOXB2 78824.391 9q21.13 forkhead box B2
GNA14 79228.369 9q21 guanine nucleotide binding protein (G protein), alpha 14
GNAQ 79525.020 9q21 guanine nucleotide binding protein (G protein), q polypeptide
PSAT1 80101.879 9q21.2 phosphoserine aminotransferase 1
TLE1 83388.420 9q21.32 transducin-like enhancer of split 1 (E(sp1) homolog, Drosophila)
RASEF 84787.137 9q21.32 RAS and EF-hand domain containing
FRMD3 85052.127 9q21.32 FERM domain containing 3
HNRNPK 85772.912 9q21.32-q21.33 heterogeneous nuclear ribonucleoprotein K
HNRPK 85772.912 9q21.32-q21.33 heterogeneous nuclear ribonucleoprotein K
RMI1 85785.457 9q21.32 RMI1, RecQ mediated genome instability 1, homolog (S. cerevisiae)
NTRK2 86473.286 9q22.1 neurotrophic tyrosine kinase, receptor, type 2
GOLM1 87830.880 9q21.33 golgi membrane protein 1
GAS1 88749.099 9q21.3-q22 growth arrest-specific 1
DAPK1 89302.576 9q34.1 death-associated protein kinase 1
CTSL1 89530.794 9q21-q22 cathepsin L1
CCRK 89771.181 9q22.1 cell cycle related kinase
SPIN1 90193.117 9q22.1-q22.3 spindlin 1
EDG3 90796.182 9q22.1-q22.2 endothelial differentiation, sphingolipid G-protein-coupled receptor, 3
S1PR3 90796.182 9q22.1-q22.2 sphingosine-1-phosphate receptor 3
SHC3 90817.889 9q22.1 SHC (Src homology 2 domain containing) transforming protein 3
CKS2 91115.933 9q22 CDC28 protein kinase regulatory subunit 2
SEMA4D 91181.972 9q22-q31 sema domain, immunoglobulin domain (Ig), transmembrane domain (TM) and short cytoplasmic domain, (semaphorin) 4D
GADD45G 91409.748 9q22.1-q22.2 growth arrest and DNA-damage-inducible, gamma
DIRAS2 92411.934 9q22.2 DIRAS family, GTP-binding RAS-like 2
SYK 92603.891 9q22 spleen tyrosine kinase
ROR2 93524.705 9q22 receptor tyrosine kinase-like orphan receptor 2
OGN 94186.070 9q22 osteoglycin
OMD 94216.359 9q22.31 osteomodulin
ECM2 94297.420 9q22.3 extracellular matrix protein 2, female organ and adipocyte specific
BICD2 94513.467 9q22.31 bicaudal D homolog 2 (Drosophila)
NINJ1 94923.592 9q22 ninjurin 1
WNK2 94987.033 9q22.3 WNK lysine deficient protein kinase 2
FBP2 96360.826 9q22.3 fructose-1,6-bisphosphatase 2
FBP1 96405.238 9q22.3 fructose-1,6-bisphosphatase 1
FANCC 96901.159 9q22.3 Fanconi anemia, complementation group C
PTCH1 97245.085 9q22.3 patched homolog 1 (Drosophila)
C9orf102 97677.721 9q22.32 RAD26L hypothetical protein
SLC35D2 98122.810 9q22.32 solute carrier family 35, member D2
CDC14B 98302.218 9q22.33 CDC14 cell division cycle 14 homolog B (S. cerevisiae)
CTSL2 98834.761 9q22.2 cathepsin L2
TMOD1 99326.258 9q22.3 tropomodulin 1
XPA 99477.012 9q22.3 xeroderma pigmentosum, complementation group A
FOXE1 99655.358 9q22 forkhead box E1 (thyroid transcription factor 2)
HEMGN 99728.896 9q22.33 hemogen
ANP32B 99785.310 9q22.32 acidic (leucine-rich) nuclear phosphoprotein 32 family, member B
NANS 99858.842 9p24.1-p23 N-acetylneuraminic acid synthase (sialic acid synthase)
GALNT12 100609.802 9q22.33 UDP-N-acetyl-alpha-D-galactosamine:polypeptide N-acetylgalactosaminyltransferase 12 (GalNAc-T12)
TGFBR1 100907.233 9q22 transforming growth factor, beta receptor I (activin A receptor type II-like kinase, 53kDa)
SEC61B 101024.391 9q22.32-q31.3 Sec61 beta subunit
NR4A3 101623.958 9q22 nuclear receptor subfamily 4, group A, member 3
TMEFF1 102275.538 9q31 transmembrane protein with EGF-like and two follistatin-like domains 1
ALDOB 103223.480 9q21.3-q22.2 aldolase B, fructose-bisphosphate
RNF20 103335.954 9q22 ring finger protein 20
PPP3R2 103393.720 9q31.1 protein phosphatase 3 (formerly 2B), regulatory subunit B, beta isoform
SMC2 105896.362 9q31.1 structural maintenance of chromosomes 2
TAL2 107464.599 9q32 T-cell acute lymphocytic leukemia 2
RAD23B 109085.365 9q31.2 RAD23 homolog B (S. cerevisiae)
KLF4 109286.956 9q31 Kruppel-like factor 4 (gut)
IKBKAP 110669.622 9q31 inhibitor of kappa light polypeptide gene enhancer in B-cells, kinase complex-associated protein
CTNNAL1 110744.672 9q31.2 catenin (cadherin-associated protein), alpha-like 1
EPB41L4B 110974.076 9q31-q32 erythrocyte membrane protein band 4.1 like 4B
PTPN3 111180.339 9q31 protein tyrosine phosphatase, non-receptor type 3
TXN 112046.131 9q31 thioredoxin
MUSK 112470.960 9q31.3-q32 muscle, skeletal, receptor tyrosine kinase
EDG2 112675.877 9q31.3 endothelial differentiation, lysophosphatidic acid G-protein-coupled receptor, 2
GNG10 113463.682 9q31.3 guanine nucleotide binding protein (G protein), gamma 10
UGCG 113699.027 9q31 UDP-glucose ceramide glucosyltransferase
POLE3 115209.342 9q33 polymerase (DNA directed), epsilon 3 (p17 subunit)
RGS3 115382.761 9q32 regulator of G-protein signaling 3
AMBP 115862.231 9q32-q33 alpha-1-microglobulin/bikunin precursor
ORM1 116125.157 9q31-q32 orosomucoid 1
ORM2 116131.890 9q32 orosomucoid 2
TNFSF15 116591.434 9q32 tumor necrosis factor (ligand) superfamily, member 15
TNFSF8 116704.945 9q33 tumor necrosis factor (ligand) superfamily, member 8
TNC 116822.626 9q33 tenascin C (hexabrachion)
DEC1 116943.918 9q32 deleted in esophageal cancer 1
PAPPA 117955.892 9q33.2 pregnancy-associated plasma protein A, pappalysin 1
ASTN2 118227.328 9q33.1 astrotactin 2
TRIM32 118489.402 9q33.1 tripartite motif-containing 32
TLR4 119506.431 9q32-q33 toll-like receptor 4
DBC1 120968.729 9q32-q33 deleted in bladder cancer 1
CDK5RAP2 122190.969 9q33.2 CDK5 regulatory subunit associated protein 2
FBXW2 122559.075 9q34 F-box and WD repeat domain containing 2
PHF19 122657.752 9q33.2 PHD finger protein 19
TRAF1 122704.493 9q33-q34 TNF receptor-associated factor 1
CEP110 122890.395 9q33-q34 centrosomal protein 110kDa
RAB14 122980.237 9q32-q34.11 RAB14, member RAS oncogene family
GSN 123070.201 9q33 gelsolin (amyloidosis, Finnish type)
GGTA1 123257.139 9q33.2-q34.11 glycoprotein, alpha-galactosyltransferase 1
DAB2IP 123544.885 9q33.1-q33.3 DAB2 interacting protein
LHX6 124004.679 9q33.2 LIM homeobox 6
PTGS1 124173.050 9q32-q33.3 prostaglandin-endoperoxide synthase 1 (prostaglandin G/H synthase and cyclooxygenase)
psMAGEA 124328.458 Xq28 melanoma antigen pseudogene, family A
NEK6 126060.070 9q33.3-q34.11 NIMA (never in mitosis gene a)-related kinase 6
NR5A1 126283.337 9q33 nuclear receptor subfamily 5, group A, member 1
NR6A1 126324.524 9q33-q34.1 nuclear receptor subfamily 6, group A, member 1
PPP6C 126951.230 9q33.3 protein phosphatase 6, catalytic subunit
HSPA5 127036.955 9q33-q34.1 heat shock 70kDa protein 5 (glucose-regulated protein, 78kDa)
MAPKAP1 127323.572 9q33.3 mitogen-activated protein kinase associated protein 1
PBX3 127549.487 9q33-q34 pre-B-cell leukemia homeobox 3
RALGPS1 128716.874 9q33.3 Ral GEF with PH domain and SH3 binding motif 1
ANGPTL2 128889.449 9q34 angiopoietin-like 2
SH2D3C 129540.419 9q34.11 SH2 domain containing 3C
CDK9 129588.158 9q34.1 cyclin-dependent kinase 9
FPGS 129604.975 9q34.1 folylpolyglutamate synthase
ENG 129617.117 9q33-q34.1 endoglin (Osler-Rendu-Weber syndrome 1)
AK1 129668.580 9q34.1 adenylate kinase 1
PIP5KL1 129723.631 9q34.11 phosphatidylinositol-4-phosphate 5-kinase-like 1
LCO 129951.171 2q14-q21 liver cancer oncogene
LCN2 129951.553 9q34 lipocalin 2
CIZ1 129968.166 9q34.1 CDKN1A interacting zinc finger protein 1
DNM1 130005.484 9q34 dynamin 1
SET 130491.682 9q34 SET translocation (myeloid leukemia-associated)
PKN3 130504.623 9q34.11 protein kinase N3
ENDOG 130620.600 9q34.1 endonuclease G
PPP2R4 130913.434 9q34 protein phosphatase 2A activator, regulatory subunit 4
IER5L 130977.652 9q34.11 immediate early response 5-like
PRRX2 131467.741 9q34.1 paired related homeobox 2
PTGES 131540.433 9q34.3 prostaglandin E synthase
C9orf78 131629.386 9q34.11 chromosome 9 open reading frame 78
FNBP1 131689.287 9q34 formin binding protein 1
ASS1 132309.915 9q34.1 argininosuccinate synthetase 1
ABL1 132579.089 9q34.1 c-abl oncogene 1, receptor tyrosine kinase
LAMC3 132874.325 9q31-q34 laminin, gamma 3
NUP214 132990.802 9q34.1 nucleoporin 214kDa
RAPGEF1 133441.979 9q34.3 Rap guanine nucleotide exchange factor (GEF) 1
SETX 134126.649 9q34.13 senataxin
TTIM1 134240.756 7p12-cen T-cell tumor invasion and metastasis 1
TTF1 134240.760 9q34.13 transcription termination factor, RNA polymerase I
DDX31 134503.071 9q34.13 DEAD (Asp-Glu-Ala-Asp) box polypeptide 31
TSC1 134776.208 9q34 tuberous sclerosis 1
GFI1B 134843.919 9q34.13 growth factor independent 1B transcription repressor
CEL 134927.186 9q34.3 carboxyl ester lipase (bile salt-stimulated lipase)
CELP 134947.746 9q34.3 carboxyl ester lipase pseudogene
RALGDS 134962.929 9q34.3 ral guanine nucleotide dissociation stimulator
ABO 135120.384 9q34.1-q34.2 ABO blood group (transferase A, alpha 1-3-N-acetylgalactosaminyltransferase; transferase B, alpha 1-3-galactosyltransferase)
RPL7A 135204.890 9q34 ribosomal protein L7a
SURF1 135208.487 9q34.2 surfeit 1
ADAMTS13 135276.941 9q34 ADAM metallopeptidase with thrombospondin type 1 motif, 13
VAV2 135616.838 9q34.1 vav 2 guanine nucleotide exchange factor
BRD3 135887.784 9q34 bromodomain containing 3
RXRA 136358.231 9q34.3 retinoid X receptor, alpha
KIAA0649 137511.469 9q34.3 KIAA0649
PAEP 137593.425 9q34 progestagen-associated endometrial protein (placental protein 14, pregnancy-associated endometrial alpha-2-globulin, alpha uterine protein)
LHX3 138227.919 9q34.3 LIM homeobox 3
GPSM1 138348.726 9q34.3 G-protein signaling modulator 1 (AGS3-like, C. elegans)
CARD9 138378.229 9q34.3 caspase recruitment domain family, member 9
SDCCAG3 138416.196 9q34.3 serologically defined colon cancer antigen 3
INPP5E 138442.893 9q34.3 inositol polyphosphate-5-phosphatase, 72 kDa
NOTCH1 138508.717 9q34.3 Notch homolog 1, translocation-associated (Drosophila)
AGPAT2 138687.416 9q34.3 1-acylglycerol-3-phosphate O-acyltransferase 2 (lysophosphatidic acid acyltransferase, beta)
C9orf86 138822.195 9q34.3 chromosome 9 open reading frame 86
EDF1 138877.054 9q34.3 endothelial differentiation-related factor 1
TRAF2 138900.786 9q34 TNF receptor-associated factor 2
PTGDS 138991.777 9q34.2-q34.3 prostaglandin D2 synthase 21kDa (brain)
NPDC1 139053.744 9q34.3 neural proliferation, differentiation and control, 1
MAN1B1 139101.200 9q34 mannosidase, alpha, class 1B, member 1
GRIN1 139153.430 9q34.3 glutamate receptor, ionotropic, N-methyl D-aspartate 1
TUBB2C 139255.532 9q34 tubulin, beta 2C
COBRA1 139269.768 9q34 cofactor of BRCA1
MAGEB17 139315.857 Xp22.2 melanoma antigen family B, 17
NOXA1 139437.668 9q34.3 NADPH oxidase activator 1
MRPL41 139566.165 9q34.3 mitochondrial ribosomal protein L41


These entries belong to a list of genes implicated in cancer which is a compilation of genes
from Atlas of Genetics and Cytogenetics and some other genes selected for their interest.


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