Circumscribed Astrocytic Gliomas

2023-07-27   Scott Ryall, PhD 

1.Brigham and Women's Hospital, Harvard Medical School, Boston , MA (USA)

Classification

Definition

The 2021 WHO guidelines for CNS tumor classification recognizes circumscribed astrocytic gliomas to acknowledge the characteristic solid growth pattern, in contrast to diffuse tumors. 1 Circumscribed astrocytic gliomas include 6 entities: i) Pilocytic astrocytoma, ii) High-grade astrocytoma with piloid features (HGAP), iii) Pleomorphic xanthoastrocytoma (PXA), iv) Subependymal giant cell astrocytoma (SEGA), v) Chordoid glioma, and vi) Astroblastoma, MN1-altered

Circumscribed Astrocytic GliomasGenetic Event(s)
Pilocytic astrocytomaThe most frequent genetic abnormalities observed in pilocytic astrocytoma are rearrangements of approximately 2-Mb at chromosome 7q34 resulting in the KIAA1549::BRAF gene fusion involving various combinations of the KIAA1549 and BRAF exons. 2-5 Infrequently, alternate BRAF fusions have been observed 4,6-8, which, like KIAA1549::BRAF, all result in loss of the N-terminal regulatory domain of BRAF, leading to dysregulated MAPK signaling. 5,9,10 Other genetic abnormalities are in genes encoding other members of the MAPK pathway and include NF1 alterations (typically germline and associated with optic pathway glioma), BRAF mutations (most often p.V600E), FGFR1 mutations, FGFR1 fusions (most often FGFR1::TACC1), NTRK1/NTRK2/NTRK3 fusions, KRAS mutations, and RAF1 fusions. 5,9,11-13
High-grade astrocytoma with piloid featuresCurrently, high-grade astrocytoma with piloid features (HGAP) can only be diagnosed via DNA methylation profiling. 14,15 Despite this, certain genetic markers suggestive of the diagnosis include CDKN2A and/or CDKN2B homozygous deletions (~80%), ATRX mutations and/or loss of expression (45%), KIAA1549::BRAF (20%), NF1 mutations or deletions (~30%), FGFR1 mutations (~20%), KRAS mutations (~3%), and BRAF mutations (~1%). 14 Total or partial loss of chromosome 19q is observed in approximately 50% of HGAP. 14,15
Pleomorphic xanthoastrocytomaAll pleomorphic xanthoastrocytoma (PXA) harbor activating genetic alterations in a MAPK pathway gene. The most frequent of these is BRAF p.V600E which is seen in up to 80% of tumors. 5,16,17 >90% of BRAF p.V600E tumors have combined homozygous deletions of CDKN2A and/or CDKN2B. 16,18-20 Less frequently, these tumors may harbor TERT promoter mutations or amplifications. 16,21 Tumors without BRAF p.V600E can harbour a wide variety of alternative alterations in the MAPK pathway including non-canonical BRAF mutations and fusions, NTRK1/NTRK2/NTRK3 fusions, and NF1 alterations. Rare alterations in SMARCB1, BCOR, ARID1A,ATRX, PTEN, FANCA, PRKDC, NOTCH2/NOTCH3/NOTCH4, and BCL6 have also been described, but their significance is, as of now, unclear. 16,19,22 DNA methylation analysis classifies PXA as its own entity, which my help alleviate diagnsotic concern in cases with ambiguous morphology or non-canonical genetics. 15
Subependymal giant cell astrocytomaSubependymal giant cell astrocytoma (SEGA) is strongly associated with tuberous sclerosis. OMIM:191100 OMIM:613254 These tumors commonly harbor biallelic inactivation of TSC1 or TSC2, with the second hit frequenty being a deletion or loss of heterozygosity. 23-25 SEGA in the absence of tuberous sclerosis have been reported, but it is unclear whether they harbor undetectable TSC1/TSC2 variants or alternative mechanisms of inactivation. 25,26 BRAF p.V600E has been infrequently reported in SEGA, although these findings are disputed. 12,25,27 DNA methylation analysis classifies SEGA as its own tumor entity. 15
Chordoid gliomaChordoid glioma are near-ubiquitously defined by p.D463H mutations in PRKCA, which is a diagnostic hallmark in these lesions. 28,29 Although the precise mechanism of this mutation remains to be fully elucidated, it has been associated with elevated levels of phosphorylated ERK, suggesting that it activates the MAPK signaling pathway. 28 Mutations in other known glioma-related genes are not seen in these tumors. DNA methylation analysis classifies chordoid glioma as its own diagnostic entity, which can be helpful in histologically ambiguous cases. 15,30
Astroblastoma, MN1-alteredAstroblastoma, MN1-altered is defined by structural rearrangements of MN1, most commonly as MN1::BEND2 fusions, although other rare fusion partners have been reported. 31-34 Typically, MN1 alterations do not co-occurr with secondary oncogenic alterations, however, a rare subset of tumors have been shown to also harbor CDKN2A homozygous deletions. 31,33 Cytogenetically, astroblastomas often harbor monosomy 16 and partial losses of 22q and X. 31-33,35 DNA methylation analysis shows that astroblastoma have a unique profile as compared to other tumors with astroblastomatous rosettes, which can be helpful in diagnostically differentiating these cases. 15

Article Bibliography

Reference NumberPubmed IDLast YearTitleAuthors
1341850762021The 2021 WHO Classification of Tumors of the Central Nervous System: a summary.Louis DN et al
2189741082008Tandem duplication producing a novel oncogenic BRAF fusion gene defines the majority of pilocytic astrocytomas.Jones DT et al
3183985032008BRAF gene duplication constitutes a mechanism of MAPK pathway activation in low-grade astrocytomas.Pfister S et al
4193738552009Activation of the ERK/MAPK pathway: a signature genetic defect in posterior fossa pilocytic astrocytomas.Forshew T et al
5322892782020Integrated Molecular and Clinical Analysis of 1,000 Pediatric Low-Grade Gliomas.Ryall S et al
6214245302011Oncogenic FAM131B-BRAF fusion resulting from 7q34 deletion comprises an alternative mechanism of MAPK pathway activation in pilocytic astrocytoma.Cin H et al
7284485142017A new GTF2I-BRAF fusion mediating MAPK pathway activation in pilocytic astrocytoma.Tomić TT et al
8291416722017A novel GIT2-BRAF fusion in pilocytic astrocytoma.Helgager J et al
9238175722013Recurrent somatic alterations of FGFR1 and NTRK2 in pilocytic astrocytoma.Jones DT et al
10235839812013Whole-genome sequencing identifies genetic alterations in pediatric low-grade gliomas.Zhang J et al
11177127322007Further evidence for a somatic KRAS mutation in a pilocytic astrocytoma.Janzarik WG et al
12212747202011Analysis of BRAF V600E mutation in 1,320 nervous system tumors reveals high mutation frequencies in pleomorphic xanthoastrocytoma, ganglioglioma and extra-cerebellar pilocytic astrocytoma.Schindler G et al
13232228492013Somatic neurofibromatosis type 1 (NF1) inactivation characterizes NF1-associated pilocytic astrocytoma.Gutmann DH et al
14295645912018Anaplastic astrocytoma with piloid features, a novel molecular class of IDH wildtype glioma with recurrent MAPK pathway, CDKN2A/B and ATRX alterations.Reinhardt A et al
15295396392018DNA methylation-based classification of central nervous system tumours.Capper D et al
16300515282019The genetic landscape of anaplastic pleomorphic xanthoastrocytoma.Phillips JJ et al
1732619301988Use of aspirin for prevention of cardiovascular disease--1981-82 to 1985-86: the Minnesota Heart Survey.Folsom AR et al
18264547672015BRAF mutation and anaplasia may be predictive factors of progression-free survival in adult pleomorphic xanthoastrocytoma.Tabouret E et al
19281813252018Recurrent copy number alterations in low-grade and anaplastic pleomorphic xanthoastrocytoma with and without BRAF V600E mutation.Vaubel RA et al
20304967962019Molecular features of pleomorphic xanthoastrocytoma.Zou H et al
21302408662018BRAF V600E, TERT, and IDH2 Mutations in Pleomorphic Xanthoastrocytoma: Observations from a Large Case-Series Study.Ma C et al
22326193052021Biology and grading of pleomorphic xanthoastrocytoma-what have we learned about it?Vaubel R et al
2394037141997Loss of tuberin in both subependymal giant cell astrocytomas and angiomyolipomas supports a two-hit model for the pathogenesis of tuberous sclerosis tumors.Henske EP et al
2490071041996Loss of tuberin from cerebral tissues with tuberous sclerosis and astrocytoma.Mizuguchi M et al
25292211452017Subependymal giant cell astrocytomas in Tuberous Sclerosis Complex have consistent TSC1/TSC2 biallelic inactivation, and no BRAF mutations.Bongaarts A et al
26259785312015Subependymal giant cell astrocytoma in the absence of tuberous sclerosis complex: case report.Beaumont TL et al
27253461652015BRAF V600E mutations are frequent in dysembryoplastic neuroepithelial tumors and subependymal giant cell astrocytomas.Lee D et al
28294761362018A recurrent kinase domain mutation in PRKCA defines chordoid glioma of the third ventricle.Goode B et al
29299152582018A recurrent point mutation in PRKCA is a hallmark of chordoid gliomas.Rosenberg S et al
30299679402018Practical implementation of DNA methylation and copy-number-based CNS tumor diagnostics: the Heidelberg experience.Capper D et al
31289606232018Multimodal molecular analysis of astroblastoma enables reclassification of most cases into more specific molecular entities.Wood MD et al
32289907082018Astroblastoma: a distinct tumor entity characterized by alterations of the X chromosome and MN1 rearrangement.Hirose T et al
33308764552019Genomic analysis demonstrates that histologically-defined astroblastomas are molecularly heterogeneous and that tumors with MN1 rearrangement exhibit the most favorable prognosis.Lehman NL et al
34311112742019MN1 rearrangement in astroblastoma: study of eight cases and review of literature.Mhatre R et al
35269194352016New Brain Tumor Entities Emerge from Molecular Classification of CNS-PNETs.Sturm D et al

Citation

Scott Ryall, PhD

Circumscribed Astrocytic Gliomas

Atlas Genet Cytogenet Oncol Haematol. 2023-07-27

Online version: http://atlasgeneticsoncology.org/solid-tumor/209195/circumscribed-astrocytic-gliomas